[{"data":1,"prerenderedAt":-1},["ShallowReactive",2],{"detail-sidebar-cat-0-en-105":3,"doc-seo-567677-105":59,"doc-detail-567677-en":130},{"code":4,"msg":5,"data":6},0,"success",[7,13,18,23,28,33,38,43,48,51,55],{"id":8,"doc_module":4,"doc_module_name":9,"category_name":10,"show_sort_weight":11,"slug":12},1,"Document","Story & Novel",90,"story-novel",{"id":14,"doc_module":4,"doc_module_name":9,"category_name":15,"show_sort_weight":16,"slug":17},2,"Literature",80,"literature",{"id":19,"doc_module":4,"doc_module_name":9,"category_name":20,"show_sort_weight":21,"slug":22},4,"Exam",70,"exam",{"id":24,"doc_module":4,"doc_module_name":9,"category_name":25,"show_sort_weight":26,"slug":27},5,"Comic",60,"comic",{"id":29,"doc_module":4,"doc_module_name":9,"category_name":30,"show_sort_weight":31,"slug":32},6,"Technology",50,"technology",{"id":34,"doc_module":4,"doc_module_name":9,"category_name":35,"show_sort_weight":36,"slug":37},7,"Healthcare",40,"healthcare",{"id":39,"doc_module":4,"doc_module_name":9,"category_name":40,"show_sort_weight":41,"slug":42},8,"Research & Report",30,"research-report",{"id":44,"doc_module":4,"doc_module_name":9,"category_name":45,"show_sort_weight":46,"slug":47},9,"Religion & Spirituality",20,"religion-spirituality",{"id":46,"doc_module":4,"doc_module_name":9,"category_name":49,"show_sort_weight":46,"slug":50},"World Cup","world-cup",{"id":52,"doc_module":4,"doc_module_name":9,"category_name":53,"show_sort_weight":52,"slug":54},10,"Lifestyle","lifestyle",{"id":56,"doc_module":4,"doc_module_name":9,"category_name":57,"show_sort_weight":24,"slug":58},19,"General","general",{"code":4,"msg":60,"data":61},"ok",{"site_id":62,"language":63,"slug":64,"title":65,"keywords":66,"description":67,"schema_data":68,"social_meta":123,"head_meta":125,"extra_data":127,"updated_unix":129},105,"en","proceedings-of-the-california-academy-of-sciences-bilateral-asymmetry-in-phallostethid-fishes-atherinomorpha","PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES - Bilateral Asymmetry in Phallostethid Fishes (Atherinomorpha)","","Bilateral asymmetry in phallostethid fishes is compared to clarify the nature of this phenomenon and its relevance for determining the homology of priapial structures. The work describes Phenacostethus trewavasae, the first phallostethine known from Borneo, from Baram River freshwater streams in Sarawak, emphasizing a distinguishing minute eye-lens. Male orientation is analyzed as sinistral or dextral, with contrasting fixed patterns reported across species.",{"@graph":69,"@context":122},[70,84,105],{"@type":71,"itemListElement":72},"BreadcrumbList",[73,77,79,82],{"item":74,"name":75,"@type":76,"position":8},"https://docshare.wps.com","Home","ListItem",{"item":78,"name":9,"@type":76,"position":14},"https://docshare.wps.com/document/",{"item":80,"name":40,"@type":76,"position":81},"https://docshare.wps.com/document/research-report/",3,{"item":83,"name":65,"@type":76,"position":19},"https://docshare.wps.com/document/proceedings-of-the-california-academy-of-sciences-bilateral-asymmetry-in-phallostethid-fishes-atherinomorpha/567677/",{"url":83,"name":65,"@type":85,"image":86,"author":91,"headline":65,"publisher":94,"fileFormat":97,"inLanguage":63,"description":67,"dateModified":98,"datePublished":99,"encodingFormat":97,"isAccessibleForFree":100,"interactionStatistic":101},"DigitalDocument",{"url":87,"@type":88,"width":89,"height":90},"https://docshare.wps.com/thumbnails/proceedings-of-the-california-academy-of-sciences-bilateral-asymmetry-in-phallostethid-fishes-atherinomorpha/567677.png","ImageObject",300,407,{"name":92,"@type":93},"Sarah ","Person",{"url":74,"name":95,"@type":96},"DocShare","Organization","application/pdf","2026-10-17","2026-10-13",true,{"@type":102,"interactionType":103,"userInteractionCount":8},"InteractionCounter",{"@type":104},"ViewAction",{"@type":106,"mainEntity":107},"FAQPage",[108,114,118],{"name":109,"@type":110,"acceptedAnswer":111},"What is the main purpose of comparing bilateral asymmetry in phallostethid fishes?","Question",{"text":112,"@type":113},"To better understand the nature of bilateral asymmetry and assess its importance for determining the homology of priapial structures.","Answer",{"name":115,"@type":110,"acceptedAnswer":116},"What new species is described, and where was it collected?",{"text":117,"@type":113},"Phenacostethus trewavasae is described from cay-and gravel-bottom freshwater streams of the Baram River in Sarawak, Malaysian Borneo.",{"name":119,"@type":110,"acceptedAnswer":120},"How are male phallostethids classified in terms of body orientation, and what patterns are reported?",{"text":121,"@type":113},"Males are termed sinistral or dextral based on whether the priapial organ is oriented toward the left or right side. The text reports both types in some species (e.g., Phallostethus dunckeri and Phenacostethus smithi), exclusively dextral males in Ph. posthon, exclusively sinistral males in P. trewavasae, and a single neostethine species with exclusively dextral males (Mirophallus mikolasi).","https://schema.org",{"og:url":83,"og:type":124,"og:title":65,"og:site_name":95,"og:description":67},"article",{"robots":126,"canonical":83},"index,follow",{"doc_id":128,"site_id":62},567677,1792251884,{"code":4,"msg":5,"data":131},{"doc_id":128,"user_id":132,"nickname":92,"user_avatar":133,"doc_module":4,"category_id":39,"category_name":40,"doc_title":65,"doc_description":67,"doc_content":134,"file_id":135,"file_url":136,"file_type":137,"file_size":138,"view_count":8,"is_deleted":4,"is_public":8,"is_downloadable":8,"audit_status":8,"page_count":139,"language":140,"language_code":63,"site_id":62,"html_lang":63,"table_of_contents":141,"faqs":142,"seo_title":143,"seo_description":67,"update_tm":144,"read_time":145},962085320529,"https://ap-avatar.wpscdn.com/davatar_9964176cb1d06d4a9deccf72a44ae3dc","# PROCEEDINGSOF THECALIFORNIA ACADEMY OF SCIENCES\n\nBILATERAL ASYMMETRY IN PHALLOSTETHID FISHES(ATHERINOMORPHA)WITH DESCRIPTION OFA NEW SPECIES FROM SARAWAK  \nBlateral asymmetry is compared among phallostethids to assess better the nature of this phenomenon,andits importance in determining the homology of priapial structares.  \n## INTRODUCTION\n\nPhallostethids are a group of some 20 knownspecies of Indo-Australian atherinomorph fishesdistinguished from all other teleosts by the pres-ence in males of the priapium,a complex,sub-cephalic copulatory organ(Regan 1913,1916).Phallostethidshave been divided into two groups(and classified traditionally in two families,Phal-lostethidae and Neostethidae,as in Roberts1971a,b)on the basis of gross differences inpriapial morphology.Rosen and Parenti(1981)and Parenti(1984)treated the entire group asone family,the Phallostethidae sensu lato,andthat convention is followed here;the two groupsare referred to as the subfamilies Phallostethinae  \nMay 6,1986  \nByLynne R.ParentiCaliforaiaAcndemy of Sciences,Golden Gate Park,San Franciaea,Califoraia 94118  \nAaSTRACT:Phenacostethas trewarsee,the finst phallostethine from Borneo,is described from cay-andgravel-bottom freshwater streams of the Baram River,Sarawak.One of the diatinguishing characteristics ofP.treinraae is a minute eye-leas.The subfamlly Phallostethinae comprises Phallostethas Regan,knownonly from a single collectiom of one spedes2.dunckeri Regan,frem the mosth of the Mar River,in Johoreon the Malay Peninsnla;and Phenacostethas Myen,known previoosly from twe species,P.smithi Myenand P.porthen Roberts,from coastal peninsular Malaysin and Thalland.  \nMale phallostethids are bilaterally aaymmetrle.The subcephalle copulatory organ,the priaplum,is orientedso that the aproctal side of the body is either the keft or right;hence,males are tenmed sinistral or dextral,respectively.Both Phallosterhas dunckeri and Phenacostethas smitki have,in about equal sumbers,malesthat are either sinistral or dextral.InP.posthon,all males are dextral,whereas,in P.trewasne,all malesare sinistral.One specles of neostethine,Mirophallus Mikolanas Herre,ls kaowa in which all males arelextral.  \nand Neostethinae.The closest living relative ofthe freshwater,brackish,and occasionally salt-water phallostethids is hypothesized to be themonotypic western Pacific marine silverside(orhardyhead)Dentatherina Patten and Ivantsoff(Parenti 1984).  \nThe subfamily Phallostethinae includes twogenera:Phallostethus Regan with one species,P.dunckeriRegan,1913,and Phenacostethus Myers,with three species,P.smithi Myers,1928,P.pos-thon Roberts,1971a,and P.trewavasae,de-scribed herein.  \nPhallostethids are found throughout coastalpeninsular Malaysia,Thailand,Borneo,the Phil-ippines,and Java.Phallostethus and Phenaco-stethus were known previously only from penin-  \nsular Malaysia and Thailand(see Roberts 1971a,fig.1).The third species of Phenacostethus,P.trewavasae,was collected from the Baram Riverin Sarawak,Malaysian Borneo.Fowler(1937)described Phenacostethus thai from a series ofnine specimens,four males and five females(Academy of Natural Sciences,Philadelphia,ANSP 51352-51360).Roberts(1971a)followedHerre(1942)in treating Phenacostethus thai asa synonym of Phenacostethus smithi,and I ten-tatively concur.  \nAll Phallostethinae and Neostethinae males arebilaterally asymmetricwith regard to orientationof the major supporting bones and other struc-tures associated with the priapium.Most femalesare bilaterally symmetric;the anus is just ante-rior to the urogenital opening along the ventralmidline,under the throat (see Regan 1913,fig.  \n2).In just one species of neostethine are bilat-erally asymmetric females found(see Parenti,inpress).  \nBilateral asymmetry is well documented infishes(see Hubbs and Hubbs 1945,and refer-ences therein,for a comprehensive review).Mostphallostethid species have both sinistral and dex-tral males in more orless equalnumbers.Roberts(1971a)described Phenacostethus pos","cbCaiqNhQi20BK5E","https://ap.wps.com/l/cbCaiqNhQi20BK5E","pdf",3472973,13,"English","# Introduction\n# Materials and Methods","[{\"question\":\"What is the main purpose of comparing bilateral asymmetry in phallostethid fishes?\",\"answer\":\"To better understand the nature of bilateral asymmetry and assess its importance for determining the homology of priapial structures.\"},{\"question\":\"What new species is described, and where was it collected?\",\"answer\":\"Phenacostethus trewavasae is described from cay-and gravel-bottom freshwater streams of the Baram River in Sarawak, Malaysian Borneo.\"},{\"question\":\"How are male phallostethids classified in terms of body orientation, and what patterns are reported?\",\"answer\":\"Males are termed sinistral or dextral based on whether the priapial organ is oriented toward the left or right side. The text reports both types in some species (e.g., Phallostethus dunckeri and Phenacostethus smithi), exclusively dextral males in Ph. posthon, exclusively sinistral males in P. trewavasae, and a single neostethine species with exclusively dextral males (Mirophallus mikolasi).\"}]","PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES - Bilateral Asymmetry in Phallostethid Fishes (Atherinomorpha) | PDF",1791867438,33]